High Quality Content by WIKIPEDIA articles! Archaic human admixture with modern Homo sapiens is suspected by some to have occurred at least twice in history: with Neanderthals, and with the population to which the Denisova hominin belonged. A minimum estimated 1% to 4% of the DNA in Eurasians, North Africans is non-modern, and shared with ancient Neanderthal DNA rather than with Sub-Saharan Africans (i.e. Yoruba and San probands). However, in August 2012, a study by scientists at the University of Cambridge has questioned this conclusion, hypothesising instead that the DNA overlap is a remnant of a common ancestor of both Neanderthals and modern humans. A minimum additional estimated 4–6% of Melanesian DNA is from the archaic Denisovan hominins from Asia. Recent DNA analysis also indicates African admixture with a now extinct archaic population.